Day length decides roughly when to go, because it is the one signal that never drifts. Weather decides the night: migrants wait, then leave in pulses when the wind turns favourable.
EstablishedSpecialists would state this without hedging. Multiple independent lines of evidence agree.
Migratory departure is primarily under photoperiodic control acting through the endogenous annual cycle, with weather — particularly wind direction and the passage of synoptic systems — determining departure within that window. Across twenty-three years of United States weather-radar data, atmospheric conditions accounted for the great majority of variation in nightly migration intensity.
- Who this applies to
- Best established in nocturnally migrating birds of the northern temperate zone, which is where both the captive work and the radar coverage are.
- Studied in
- Aves
Why we rate it this way, and what the caveats are
Two independent lines: captive birds held on manipulated day lengths shift their migratory restlessness, and continental radar shows departure intensity tracking weather with forecastable skill.
How far it can be extended
Photoperiodic control of annual cycles is general across temperate vertebrates; the radar evidence for weather-driven timing is continental but from one network.
Caveats
- Radar measures departure intensity, not whether any individual left on its optimal night.
- Photoperiod is the dominant cue in temperate migrants; tropical and irruptive migrants are timed differently and are much less studied.
- Day length does not drift and springs do — so an unchanged photoperiodic response can now deliver a bird after the food peak it evolved to meet.
Still unanswered
- How much individual flexibility birds have to adjust departure within the photoperiodic window, which determines how fast populations can track a shifting spring.
Last reviewed 2026-09-03
The evidence (3 studies)
Supports · primary
A continental system for forecasting bird migration
Van Doren and Horton, 2018 · Science
Twenty-three years of radar showing that how much is aloft on a night is largely predictable from the weather.
Supports · supporting
Genetic Basis of Migratory Behavior in European Warblers
Berthold and Querner, 1981 · Science
The endogenous side: timing and amount of migratory restlessness persist in captivity and are heritable.
Supports · supporting
Long-distance migration: evolution and determinants
Alerstam et al., 2003 · Oikos
Places timing among the constraints that shape whether and when migration pays.