Study
Nonlinear effects of group size on the success of wolves hunting elk
MacNulty, Daniel R.; Smith, Douglas W.; Mech, L. David; Vucetich, John A.; Packer, Craig
Behavioral Ecology, 2012
Directly observed wolf hunts of elk in Yellowstone across several winters, recording group size and outcome, and modelled how the probability of a kill changed as the number of hunting wolves increased.
- Studied in
- Canis lupus, Cervus canadensis
- Sample size
- directly observed wolf hunts in Yellowstone over multiple winters
Hunting success rose sharply from one to about four wolves and then levelled off, with additional wolves adding little. The plateau is attributed to individuals withholding effort once enough others are engaged, since the benefit of a kill is shared regardless.
What it means
The authors' reading, and ours. Where they differ, that difference is the point.
How the authors put it
Larger hunting groups are not proportionally more effective, and the flattening reflects a free-riding problem rather than a physical limit.
How NatureHQ reads it
A measured answer to a question usually settled by assertion. It is also the clearest case NatureHQ holds of a group behaviour that is not straightforwardly cooperative: the animals that hang back are not failing at cooperation, they are responding to the fact that they will eat either way.
- One population, hunting one principal prey species, in winter conditions.
- Effort withheld is inferred from the shape of the success curve rather than measured for each individual.
Counting hunts until the curve appeared
Do more wolves make a better hunting group, and does that keep being true?
What this study is used for on NatureHQ
One study can inform several subjects. Here is everywhere this one is cited.
Predation
A hunt is a sequence — detect, approach, attack, subdue, keep — and it can fail at every stage. Where success has been measured properly, most attempts end without a kill.
Cooperative hunting
Several animals hunting the same prey at once is not cooperation. Cooperation means individuals taking roles that depend on each other — and that is a harder thing to show, demonstrated in far fewer species.
Cooperative hunting
Wolves hunting elk get better as the group grows to about four, and then stop improving. Extra wolves add little, apparently because an animal that hangs back still eats.
Published by: University press journalsPublished in a university press journal.
Checked against Crossref on 2026-09-28, and they agree on the title, the authors and the year.
NatureHQ summarises research in its own words and does not reproduce published text. Reviewed 2026-09-03.