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Study

Diversity in tropical rain forests and coral reefs

Connell, Joseph H.

Science, 1978

Peer-reviewednarrative reviewnot applicable

Proposes that high diversity in forests and reefs is maintained by disturbance at intermediate frequency and intensity, which prevents competitive exclusion without eliminating species that recover slowly.

Studied in
Plantae, Anthozoa
Sample size
synthesis across forest and reef diversity studies

Diversity was argued to peak at intermediate levels of disturbance, with low disturbance permitting dominance by superior competitors and high disturbance permitting only rapid colonisers.

What it means

The authors' reading, and ours. Where they differ, that difference is the point.

How the authors put it

Neither equilibrium nor constant catastrophe maintains diversity; intermediate disturbance does.

How NatureHQ reads it

One of the most cited hypotheses in ecology, and one whose empirical record is far more mixed than its prominence suggests. The reasoning is appealing and the hump-shaped curve is memorable, and neither is evidence. NatureHQ carries it alongside the argument that it should be abandoned, because a reader taught the curve as a finding has been taught something the literature does not support.

  • A hypothesis rather than a test, and the predicted pattern is found in a minority of studies that look for it.
  • The terms intermediate, frequency and intensity are not defined in a way that makes the prediction sharp.

What this study is used for on NatureHQ

One study can inform several subjects. Here is everywhere this one is cited.

Published by: American Association for the Advancement of SciencePublished in Science or Science Advances (AAAS).

doi.org/10.1126/science.199.4335.1302

Checked against Crossref on 2026-09-28, and they agree on the title, the authors and the year.

NatureHQ summarises research in its own words and does not reproduce published text. Reviewed 2026-09-04.