Almost everything said about "birdsong" applies only to song, which is a long learned performance — not to calls, which are short, largely innate and used by both sexes all year.
EstablishedSpecialists would state this without hedging. Multiple independent lines of evidence agree.
Avian vocalisations are conventionally divided into song — structurally complex, typically learned, usually produced in reproductive and territorial contexts — and calls, which are short, structurally simple, largely innate and used year-round for alarm, contact, begging and flight coordination. The developmental and functional literatures for the two categories are largely separate.
- Who this applies to
- A distinction used across birds, drawn most sharply in songbirds and blurrier in some other groups.
- Studied in
- Aves
Why we rate it this way, and what the caveats are
The categories are a convention rather than a discovery, and the substantive part — that song is typically learned and most calls are not — is supported by isolation and deafening experiments across many species.
How far it can be extended
The song/call division is applied throughout the ornithological literature and the developmental difference — song learned, most calls not — has been demonstrated separately in many songbird species.
Caveats
- The boundary is a convention and there are intermediate vocalisations that sit awkwardly on either side of it.
- Some calls are partly learned, and in a few species call learning is substantial.
- Structural simplicity is not functional simplicity — alarm calls carry information about predator type and urgency.
Still unanswered
- How much of the vocal repertoire outside song is learned in species that have not been tested, which is most of them.
Last reviewed 2026-09-02
The evidence (3 studies)
Supports · primary
Bird Song: Biological Themes and Variations
Catchpole and Slater, 2008 · Cambridge University Press
The standard synthesis, which sets out the distinction and the separate literatures behind each side of it.
Supports · supporting
Thorpe, 1958 · Ibis
Isolation removes the learned structure of song while leaving the bird still vocalising, which is the developmental difference made visible.
Qualifies · supporting
Experimental evidence for compositional syntax in bird calls
Suzuki et al., 2016 · Nature Communications
Shows that calls are not the simple side of the division in every respect: tit calls combine according to an order rule that changes the response.